Box No 55 mk I doesn't follow the normal British naming convention of a letter followed by numbers.
pic courtesy
Bonnex
My take on the naming conventions is that if it is too large to go inside something else (ie is a primary package) it follows the above nomenclature type otherwise it is a box/tube/cylinder/case and is contained in a Primary Package.
Examples
H.3 mk I (containing Carton No 6) (primary package)
Box 511 mk I (internal box)
Case, Cartridge No. 65 mk III (internal case)
There are no examples of "Box xx" etc given in any of my references.
Your Box No 55 mk I has the appearance of a Primary Package (my term, not the official one) by dint of its apparent size and sturdy construction.
I can't help you any further at the moment.